RT Journal Article T1 A new species and three hybrids in the ferns from Cocos Island, Puntarenas, Costa Rica T2 Una nueva especie y tres híbridos en helechos para la Isla del Coco, Puntarenas, Costa Rica A1 Rojas-Alvarado, Alexander Francisco K1 Helechos - Costa Rica AB A new species and three new hybrids of ferns from the Cocos Island (CostaRica) are described here: Cyathea × robusta A. Rojas, Elaphoglossum ×intermedium A. Rojas, Nephrolepis cocosensis A. Rojas and Tectaria ×epilithica A. Rojas. The new species differs from Nephrolepis rivularis (Vahl)Mett. ex Krug by its terrestrial habitat, smaller fronds, smaller pinnae thatare perpendicular to rachis and lobed to bilobed distally, pale brown rachisscales with dark brown insertion points, blade scales 0.2–0.4 mm in diameter,sometimes reduced to a resinous dots, sori submarginal, frequently in theinternal lobe of each segments, and indusia cordiform and basally attached,pale brown with dark brown point of attachment. The first hybrid is intermediatebetween Cyathea alfonsiana L.D. Gómez and C. notabilis Domin, in trunk tall,size and form of rhizome and stipe scales, fronds size, pairs per segment anddivision degree of veins and paraphyses size, but has bigger fronds that bothprogenitors, this character is the basis for its specific epithet. Elaphoglossumx intermedium is a hybrid between E. alvaradoanum A. Rojas and E. crinitum (L.) Christ and is characterized by brown stipe and blade scales, cuneate bladebase an obtuse blade apex, and veins partially and irregularly anastomosing.Tectaria x epilithica has intermediate characters between T. incisa Cav. andT. mexicana (Fée) C.V. Morton, this hybrid is easy to recognize because has1-pinnate-pinnatifid blade and possesses rhizome scales that are intermediatein size compared to those of its parents. PB Universidad de Málaga. Servicio de Publicaciones e Intercambio Científico SN 0210-9506 YR 2017 FD 2017 LK https://hdl.handle.net/10630/15812 UL https://hdl.handle.net/10630/15812 LA eng NO Acta Botánica Malacitana, Vol. 42, nº 1, 2017, págs. 91-99 DS RIUMA. Repositorio Institucional de la Universidad de Málaga RD 19 ene 2026