RT Journal Article T1 Metapopulation Patterns of Iberian Butterflies Revealed by Fuzzy Logic A1 Pulido-Pastor, Antonio A1 Márquez-Moya, Ana Luz A1 Guerrero, José Carlos A1 García-Barros, Enrique A1 Real-Giménez, Raimundo K1 Mariposas AB Metapopulation theory considers that the populations of many species are fragmentedinto patches connected by the migration of individuals through an interterritorial matrix. We appliedfuzzy set theory and environmental favorability (F) functions to reveal the metapopulationalstructure of the 222 butterfly species in the Iberian Peninsula. We used the sets of contiguous gridcells with high favorability (F ≥ 0.8), to identify the favorable patches for each species. We superimposedthe known occurrence data to reveal the occupied and empty favorable patches, as unoccupiedpatches are functional in a metapopulation dynamics analysis. We analyzed the connectivitybetween patches of each metapopulation by focusing on the territory of intermediate and lowfavorability for the species (F < 0.8). The friction that each cell opposes to the passage of individualswas computed as 1‐F. We used the r.cost function of QGIS to calculate the cost of reaching each cellfrom a favorable patch. The inverse of the cost was computed as connectivity. Only 126 species canbe considered to have a metapopulation structure. These metapopulation structures are part of thedark biodiversity of butterflies because their identification is not evident from the observation ofthe occurrence data but was revealed using favorability functions PB MDPI YR 2021 FD 2021-04 LK https://hdl.handle.net/10630/21632 UL https://hdl.handle.net/10630/21632 LA eng NO Pulido‐Pastor, A.; Márquez, A.L.; Guerrero, J.C.; García‐Barros, E. Metapopulation Patterns of Iberian Butterflies Revealed by Fuzzy Logic. Insects 2021, 12, 392. https://doi.org/10.3390/ insects12050392 NO Funding for open access charge: Universidad de Málaga / CBUA DS RIUMA. Repositorio Institucional de la Universidad de Málaga RD 22 ene 2026